Araneomorphae

Araneomorph spiders
Temporal range: Triassic–present
Nephila inaurata (Nephilidae)
Pholcus phalangioides female with eggsac (Pholcidae)
Scientific classification Edit this classification
Kingdom: Animalia
Phylum: Arthropoda
Subphylum: Chelicerata
Class: Arachnida
Order: Araneae
Suborder: Opisthothelae
Infraorder: Araneomorphae
Subdivisions
Diversity
95 families

The Araneomorphae (also called the Labidognatha or "true spiders"[1]) are an infraorder of spiders. They are distinguishable by chelicerae (fangs) that point diagonally forward and cross in a pinching action, in contrast to those of Mygalomorphae (tarantulas and their close kin), which point straight down. Araneomorphs comprise the vast majority (about 92%) of living spiders.

Distinguishing characteristics

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Most spider species are Araneomorphae, which have fangs that face towards each other, increasing the orientations that they can employ during prey-capture. They have fewer book lungs (when present) – usually one pair – and the females typically live one year.

The Mygalomorphae have fangs that face towards the ground, and which are parallel to the long axis of the spider's body, thus they have only one orientation they can employ during prey capture. They have two pairs of book lungs, and the females often live many years.[2]

Spiders included

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Almost all of the familiar spiders are included in the Araneomorphae group, one major exception being the tarantulas. There are a few other Mygalomorphae species that live around homes or gardens, but they typically are relatively small and not easily noticed.

The Araneomorphae, to the contrary, include the weavers of spiral webs; the cobweb spiders that live in the corners of rooms, and between windows and screens; the crab spiders that lurk on the surfaces of flowers in gardens; the jumping spiders that are visible hunting on surfaces; the wolf spiders that carpet hunting sites in sunny spots; and the large huntsman spiders.

Systematics

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In older schemes, the Araneomorphae were divided into two lineages, the Hypochilae (containing only the family Hypochilidae), and the Neocribellatae. The Neocribellatae were in turn divided into the Austrochiloidea, and the two series Haplogynae and Entelogynae, each containing several superfamilies. Molecular phylogenetic studies have shown that the haplogynes in particular are not a monophyletic group. A 2020 study suggested the relationships among the major groups were as shown in the following cladogram.[3]

The blue bar to the right shows the former Haplogynae in the sense of Coddington (2005).[4]

Table of families

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Key
Genera 1 ≥2 ≥10 ≥100
Species 1–9 ≥10 ≥100 ≥1000
Spider families[note 1]
Family Genera Species Common name Example
Agelenidae 100 1481 araneomorph funnel-web spiders Hobo spider (Eratigena agrestis)
Amaurobiidae 27 208 tangled nest spiders Callobius claustrarius
Anapidae 59 233 Holarchaea novaeseelandiae
Ancylometidae 1 10
Anyphaenidae 59 654 anyphaenid sac spiders Hibana velox (yellow ghost spider)
Araneidae 197 3157 orb-weaver spiders Zygiella x-notata
Archaeidae 6 93 pelican spiders Madagascarchaea gracilicollis
Archoleptonetidae 2 8 Archoleptoneta gertschi
Argyronetidae 12 74
Arkyidae 2 38
Austrochilidae 2 9 Tasmanian cave spider (Hickmania troglodytes)
Caponiidae 21 157 Diploglena capensis
Cheiracanthiidae 15 385 Cheiracanthium mildei'
Cicurinidae 4 183
Cithaeronidae 2 10
Clubionidae 18 681 sac spiders Clubiona trivialis
Corinnidae 76 892 dark sac spiders Castianeira sp.
Ctenidae 48 606 wandering spiders Phoneutria fera
Cyatholipidae 23 58
Cybaeidae 24 308 Cryphoeca silvicola
Cycloctenidae 9 81
Deinopidae 3 71 net-casting spiders Asianopis subrufa (rufous net-casting spider)
Desidae 63 323 intertidal spiders Phryganoporus candidus
Dictynidae 45 339 Nigma walckenaeri
Diguetidae 2 16 coneweb spiders
Dolomedidae 7 128
Drymusidae 2 19 false violin spiders
Dysderidae 24 666 woodlouse hunter spiders Woodlouse spider (Dysdera crocata)
Eresidae 9 120 velvet spiders Eresus sandaliatus
Filistatidae 18 192 crevice weavers Southern house spider (Kukulcania hibernalis)
Fonteferreidae 1 1
Gallieniellidae 5 41
Gnaphosidae 154 2498 flat-bellied ground spiders Drassodes cupreus
Gradungulidae 8 20 large-clawed spiders Progradungula carraiensis (Carrai cave spider)
Hahniidae 29 244 dwarf sheet spiders
Hersiliidae 16 189 tree trunk spiders Hersilia savignyi
Homalonychidae 1 2
Huttoniidae 1 1 Huttonia palpimanoides
Hypochilidae 2 33 lampshade spiders Hypochilus thorelli
Lamponidae 23 192 White-tailed spider (Lampona spp.)
Lathyidae 10 58
Leptonetidae 22 400 Tooth Cave spider (Tayshaneta myopica)
Linyphiidae 640 4965 dwarf / money spiders Linyphia triangularis
Liocranidae 35 358 liocranid sac spiders
Lycosidae 140 2510 wolf spiders Lycosa tarantula
Macrobunidae 27 95
Malkaridae 13 57 shield spiders
Mecysmaucheniidae 7 25
Megadictynidae 2 2
Mimetidae 8 166 pirate spiders Oarces reticulatus
Miturgidae 33 191 long-legged sac spiders
Myrmecicultoridae 1 2
Mysmenidae 17 188 spurred orb-weavers
Nesticidae 16 291 cave cobweb spiders Nesticella marapu
Nicodamidae 7 27
Ochyroceratidae 9 184 midget ground weavers Theotima minutissima
Oecobiidae 7 134 disc web spiders Oecobius navus
Oonopidae 115 1983 dwarf hunting spiders Oonops domesticus
Orsolobidae 30 190
Oxyopidae 10 448 lynx spiders Peucetia viridans (green lynx spider)
Pacullidae 4 38
Palpimanidae 20 182 palp-footed spiders
Penestomidae 1 9
Periegopidae 1 3
Philodromidae 31 529 philodromid crab spiders Philodromus dispar
Pholcidae 97 2060 daddy long-legs spiders Pholcus phalangioides
Phrurolithidae 27 420
Physoglenidae 13 72
Phyxelididae 14 68
Pimoidae 2 87 Pimoa cthulhu
Pisauridae 45 235 nursery web spiders Pisaura mirabilis
Plectreuridae 2 32
Prodidomidae 24 196
Psechridae 2 64
Psilodercidae 11 225
Salticidae 695 6950 jumping spiders Zebra spider (Salticus scenicus)
Scytodidae 4 262 spitting spiders Scytodes thoracica
Segestriidae 5 181 tubeweb spiders Segestria florentina
Selenopidae 9 282 wall spiders Selenops radiatus
Senoculidae 1 31
Sicariidae 3 177 recluse spiders Brown recluse (Loxosceles reclusa)
Sparassidae 99 1551 huntsman spiders Delena cancerides (Avondale spider)
Stenochilidae 2 13
Stiphidiidae 20 125 Tartarus mullamullangensis
Symphytognathidae 10 108 dwarf orb-weavers Patu digua
Synaphridae 3 13
Synotaxidae 5 40
Telemidae 16 113 long-legged cave spiders
Tetrablemmidae 27 153 armored spiders
Tetragnathidae 45 996 long jawed orb-weavers Leucauge venusta (orchard spider)
Theridiidae 138 2619 cobweb spiders Redback spider (Latrodectus hasselti)
Theridiosomatidae 24 180 ray spiders Theridiosoma gemmosum
Thomisidae 172 2194 crab spiders Misumena vatia (goldenrod crab spider)
Titanoecidae 5 67 Goeldia obscura
Toxopidae 14 82
Trachelidae 29 317
Trachycosmidae 20 148
Trechaleidae 18 136
Trochanteriidae 6 51
Trogloraptoridae 1 1 Trogloraptor marchingtoni
Udubidae 6 58
Uloboridae 20 284 hackled orb-weavers Uloborus walckenaerius
Viridasiidae 3 14
Xenoctenidae 4 33
Zodariidae 90 1325 Zodarion germanicum
Zoropsidae 28 186 Zoropsis spinimana

Extinct families

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Notes

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  1. ^ Unless otherwise shown, currently accepted families and counts based on the World Spider Catalog version 26 as of 23 January 2026[update].[5] In the World Spider Catalog, "species" counts may include subspecies. Assignment to sub- and infraorders based on Coddington (2005, p. 20) (when given there).

References

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Citations

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  1. ^ Coddington, J.A. (2005). "Phylogeny and Classification of Spiders" (PDF). In Ubick, D.; Paquin, P.; Cushing, P.E.; Roth, V. (eds.). Spiders of North America: an identification manual. American Arachnological Society. pp. 18–24. ISBN 978-0-9771439-0-0. Archived from the original (PDF) on 9 December 2008. Retrieved 12 October 2008.
  2. ^ "ABOUT SPIDERS". www.publish.csiro.au. Retrieved 2022-01-07.
  3. ^ Magalhaes, Ivan L. F.; Azevedo, Guilherme H. F.; Michalik, Peter; Ramírez, Martín J. (February 2020). "The fossil record of spiders revisited: implications for calibrating trees and evidence for a major faunal turnover since the Mesozoic". Biological Reviews. 95 (1): 184–217. doi:10.1111/brv.12559. ISSN 1464-7931. PMID 31713947. S2CID 207937170.
  4. ^ Coddington, Jonathan A. (2005). "Phylogeny and classification of spiders" (PDF). In Ubick, D.; Paquin, P.; Cushing, P.E. & Roth, V. (eds.). Spiders of North America: an identification manual. American Arachnological Society. pp. 18–24. Retrieved 2015-09-24.
  5. ^ World Spider Catalog (2026), Families.


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