List of spalacids

Gray blind mole-ratGreater blind mole-rat (Spalax microphthalmus)

Spalacidae is a family of mammals in the order Rodentia and part of the Myomorpha suborder. Members of this family are called spalacids, and include blind mole-rats, bamboo rats, mole-rats, and zokors. They are found in Asia, eastern Africa, and eastern Europe, primarily in forests, shrublands, and grasslands, though some species can be found in deserts or savannas. They range in size from the Middle East blind mole-rat, at 13 cm (5 in) with no tail, to the large bamboo rat, at 48 cm (19 in) plus a 20 cm (8 in) tail. Spalacids are primarily herbivores, and eat roots, bulbs, tubers, grass, and seeds, with some species also eating insects. Few spalacids have population estimates, but four species—the giant root-rat, Mehely's blind mole-rat, Podolsk blind mole-rat, and sandy blind mole-rat—are categorized as endangered, while the Oltenia blind mole-rat is categorized as critically endangered.

The 23 extant species of Spalacidae are divided into 7 genera, divided into 3 subfamilies. Myospalacinae contains 6 species of zokors in 2 genera, Rhizomyinae contains 6 species of bamboo rats and mole-rats in 3 genera, and Spalacinae contains 11 species of blind mole-rats in 2 genera. Several extinct prehistoric spalacid species have been discovered, though due to ongoing research and discoveries, the exact number and categorization is not fixed.[1]

Conventions

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IUCN Red List categories
Conservation status
 EX Extinct (0 species)
 EW Extinct in the wild (0 species)
 CR Critically endangered (1 species)
 EN Endangered (4 species)
 VU Vulnerable (1 species)
 NT Near threatened (0 species)
 LC Least concern (17 species)

The author citation for the species or genus is given after the scientific name; parentheses around the author citation indicate that this was not the original taxonomic placement. Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the spalacid's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.

Classification

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Spalacidae is a family consisting of 23 extant species in 7 genera. These genera are divided into four subfamilies: Myospalacinae, containing 6 species of in 2 genera; Rhizomyinae, containing 6 species in 3 genera; and Spalacinae, containing 11 species in 2 genera. This does not include hybrid species or extinct prehistoric species.

Family Spalacidae

Spalacids

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The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists.[3]

Subfamily Myospalacinae

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Genus EospalaxAllen, 1938 – three species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Chinese zokor

Gray zokor in a cage

E. fontanierii
(H. Milne-Edwards, 1867)
Central China Size: 16–24 cm (6–9 in) long, plus 3–7 cm (1–3 in) tail[4]

Habitat: Shrubland and grassland[5]

Diet: Roots and grains[6]
 LC 


Unknown Unknown[5]

Rothschild's zokor E. rothschildi
(Thomas, 1911)
Central China Size: 14–18 cm (6–7 in) long, plus 2–4 cm (1–2 in) tail[4]

Habitat: Forest, shrubland, and grassland[7]

Diet: Roots and grains[6]
 LC 


Unknown Unknown[7]

Smith's zokor E. smithii
(Thomas, 1911)
Central China Size: 16–25 cm (6–10 in) long, plus 3–4 cm (1–2 in) tail[4]

Habitat: Grassland[8]

Diet: Roots and grains[6]
 LC 


Unknown Unknown[8]

Genus MyospalaxLaxmann, 1769 – three species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
False zokor

Taxidermied brown zokor

M. aspalax
(Pallas, 1776)
Mongolia, southern Russia, and northern China Size: 16–21 cm (6–8 in) long, plus 3–7 cm (1–3 in) tail[4]

Habitat: Forest and shrubland[9]

Diet: Roots and grains[6]
 LC 


Unknown Unknown[9]

Siberian zokor

Brown zokor

M. myospalax
(Laxmann, 1773)
Kazakhstan, southern Russia, and northern China
Map of range
Size: 20–27 cm (8–11 in) long, plus about 5 cm (2 in) tail[4]

Habitat: Grassland[10]

Diet: Roots and grains[6]
 LC 


Unknown Unknown[10]

Transbaikal zokor

Taxidermied brown zokor

M. psilurus
(H. Milne-Edwards, 1874)
Mongolia, southern Russia, and northern China Size: 20–27 cm (8–11 in) long, plus 3–6 cm (1–2 in) tail[4]

Habitat: Shrubland, grassland, and desert[11]

Diet: Roots and grains[6]
 LC 


Unknown Population steady[11]

Subfamily Rhizomyinae

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Genus CannomysThomas, 1915 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Lesser bamboo rat

Brown bamboo rat

C. badius
(Hodgson, 1841)
Southeastern Asia
Map of range
Size: 14–26 cm (6–10 in) long, plus 4–10 cm (2–4 in) tail[12]

Habitat: Forest[13]

Diet: Shrubs, shoots, and roots[14]
 LC 


Unknown Unknown[13]

Genus RhizomysGray, 1831 – three species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Chinese bamboo rat

Taxidermied brown bamboo rat

R. sinensis
Gray, 1831
China, Myanmar, and Vietnam
Map of range
Size: 22–45 cm (9–18 in) long, plus 5–10 cm (2–4 in) tail[4]

Habitat: Forest[15]

Diet: Bamboo roots, as well as grass, seeds, and fruit[16]
 LC 


Unknown Unknown[15]

Hoary bamboo rat

Taxidermied brown bamboo rat

R. pruinosus
Blyth, 1851
India and southeastern Asia
Map of range
Size: 24–35 cm (9–14 in) long, plus 9–13 cm (4–5 in) tail[4]

Habitat: Forest and grassland[17]

Diet: Bamboo roots, as well as grass, seeds, and fruit[16]
 LC 


Unknown Population steady[17]

Large bamboo rat

Taxidermied brown bamboo rat

R. sumatrensis
(Raffles, 1821)
Southeastern Asia
Map of range
Size: 26–48 cm (10–19 in) long, plus 10–20 cm (4–8 in) tail[4]

Habitat: Forest[18]

Diet: Bamboo roots, as well as grass, seeds, and fruit[16]
 LC 


Unknown Unknown[18]

Genus TachyoryctesRüppell, 1835 – two species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Giant root-rat

Brown mole-rat

T. macrocephalus
Rüppell, 1842
Ethiopia Size: 22–31 cm (9–12 in) long, plus 4–7 cm (2–3 in) tail[12]

Habitat: Grassland[19]

Diet: Roots, rhizomes, tubers, bulbs, and corms, as well as grass and legumes[20]
 EN 


Unknown Population declining[19]

Northeast African mole-rat

Brown mole-rat

T. splendens
(Rüppell, 1836)
Eastern Africa Size: 15–27 cm (6–11 in) long, plus 4–10 cm (2–4 in) tail[12]

Habitat: Forest, savanna, shrubland, and grassland[21]

Diet: Roots, rhizomes, tubers, bulbs, and corms, as well as grass and legumes[20]
 LC 


Unknown Population steady[21]

Subfamily Spalacinae

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Genus NannospalaxPalmer, 1903 – three species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Anatolian blind mole-rat

Black and white drawing of mole-rats

N. xanthodon
(Nordmann, 1840)
Western Asia Size: 14–25 cm (6–10 in) long, with no tail[12]

Habitat: Grassland[22]

Diet: Roots, tubers, acorns, plant stems, and other plant parts[23]
 LC 


Unknown Unknown[22]

Lesser blind mole-rat

Gray mole-rat

N. leucodon
(Nordmann, 1840)
Southeastern Europe
Map of range
Size: 15–24 cm (6–9 in) long, with no tail[12]

Habitat: Grassland[24]

Diet: Roots, tubers, acorns, plant stems, and other plant parts[23]
 LC 


Unknown Unknown[24]

Middle East blind mole-rat

Gray mole-rat

N. ehrenbergi
(Nehring, 1898)
Middle East and northeastern Africa Size: 13–22 cm (5–9 in) long, with no tail[12]

Habitat: Shrubland, grassland, and desert[25]

Diet: Roots, tubers, acorns, plant stems, and other plant parts[23]
 LC 


Unknown Population declining[25]

Genus SpalaxGüldenstädt, 1770 – eight species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Bukovina blind mole-rat S. graecus
Nehring, 1898
Romania and southwestern Ukraine Size: 22–28 cm (9–11 in) long, with no tail[12]

Habitat: Grassland[26]

Diet: Roots, bulbs, tubers, grass, and seeds, as well as insects[23]
 VU 


Unknown Unknown[26]

Giant blind mole-rat

Brown mole-rat

S. giganteus
Nehring, 1898
Southwestern Russia Size: 25–35 cm (10–14 in) long, with no tail[12]

Habitat: Shrubland and grassland[27]

Diet: Roots, bulbs, tubers, grass, and seeds, as well as insects[23]
 LC 


Unknown Population steady[27]

Greater blind mole-rat

Gray mole-rat

S. microphthalmus
Güldenstädt, 1770
Ukraine and southwestern Russia Size: 19–31 cm (7–12 in) long, with no tail[12]

Habitat: Shrubland and grassland[28]

Diet: Roots, bulbs, tubers, grass, and seeds, as well as insects[23]
 LC 


Unknown Population steady[28]

Kazakhstan blind mole-rat S. uralensis
Tiflov & Usov, 1939
Kazakhstan Size: About 31 cm (12 in) long, with no tail[12]

Habitat: Grassland[29]

Diet: Roots, bulbs, tubers, grass, and seeds, as well as insects[23]
 LC 


Unknown Unknown[29]

Mehely's blind mole-rat S. antiquus
Méhely, 1909
Romania Size: Unknown[12]

Habitat: Grassland[30]

Diet: Roots, bulbs, tubers, grass, and seeds, as well as insects[23]
 EN 


3,500–3,800 Population declining[30]

Oltenia blind mole-rat S. istricus
Méhely, 1909
Romania Size: About 24 cm (9 in) long, with no tail[12]

Habitat: Grassland[31]

Diet: Roots, bulbs, tubers, grass, and seeds, as well as insects[23]
 CR 


Unknown Unknown[31]

Podolsk blind mole-rat

Black and white drawing of mole-rat

S. zemni
Erxleben, 1777
Ukraine Size: 20–31 cm (8–12 in) long, with no tail[12]

Habitat: Forest and grassland[32]

Diet: Roots, bulbs, tubers, grass, and seeds, as well as insects[23]
 EN 


Unknown Unknown[32]

Sandy blind mole-rat

Coin with mole-rat on it

S. arenarius
Reshetnik, 1939
Ukraine
Map of range
Size: 19–27 cm (7–11 in) long, with no tail[12]

Habitat: Grassland[33]

Diet: Roots, bulbs, tubers, grass, and seeds, as well as insects[23]
 EN 


Unknown Unknown[33]

References

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  1. ^ "Family Spalacidae Gray 1821 (mouse)". Paleobiology Database. University of Wisconsin–Madison. Archived from the original on May 16, 2025. Retrieved September 18, 2025.
  2. ^ Fabre, P.-H.; Hautier, L.; Dimitrov, D.; Douzery, E. J. P. (2012). "A glimpse on the pattern of rodent diversification: a phylogenetic approach". BMC Evolutionary Biology. 12 (1): 88. Bibcode:2012BMCEE..12...88F. doi:10.1186/1471-2148-12-88. PMC 3532383. PMID 22697210.
  3. ^ Wilson; Reeder, pp. 907–925
  4. ^ a b c d e f g h i Chernasky; Motis; Burgin, pp. 210–211
  5. ^ a b Smith, A. T.; Johnston, C. H. (2017) [errata version of 2016 assessment]. "Eospalax fontanierii". IUCN Red List of Threatened Species. 2016 e.T14118A115120816. doi:10.2305/IUCN.UK.2016-3.RLTS.T14118A22277700.en.
  6. ^ a b c d e f Nowak, p. 1429
  7. ^ a b Smith, A. T.; Johnston, C. H. (2017) [errata version of 2016 assessment]. "Eospalax rothschildi". IUCN Red List of Threatened Species. 2016 e.T14121A115121170. doi:10.2305/IUCN.UK.2016-3.RLTS.T14121A22277586.en.
  8. ^ a b Smith, A. T.; Johnston, C. H. (2017) [errata version of 2016 assessment]. "Eospalax smithii". IUCN Red List of Threatened Species. 2016 e.T14122A115121321. doi:10.2305/IUCN.UK.2016-3.RLTS.T14122A22277483.en.
  9. ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Myospalax aspalax". IUCN Red List of Threatened Species. 2016 e.T14116A115120685. doi:10.2305/IUCN.UK.2016-3.RLTS.T14116A22277071.en.
  10. ^ a b Cassola, F. (2016). "Myospalax myospalax". IUCN Red List of Threatened Species. 2016 e.T14119A22277335. doi:10.2305/IUCN.UK.2016-2.RLTS.T14119A22277335.en.
  11. ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Myospalax psilurus". IUCN Red List of Threatened Species. 2016 e.T14120A115121026. doi:10.2305/IUCN.UK.2016-3.RLTS.T14120A22277214.en.
  12. ^ a b c d e f g h i j k l m n Chernasky; Motis; Burgin, pp. 212–213
  13. ^ a b Aplin, K.; Lunde, D.; Musser, G.; Frost, A.; Molur, S. (2017) [errata version of 2016 assessment]. "Cannomys badius". IUCN Red List of Threatened Species. 2016 e.T3759A115066803. doi:10.2305/IUCN.UK.2016-3.RLTS.T3759A22274794.en.
  14. ^ Nowak, p. 1443
  15. ^ a b Lunde, D.; Aplin, K.; Musser, G. (2017) [errata version of 2016 assessment]. "Rhizomys sinensis". IUCN Red List of Threatened Species. 2016 e.T19646A115152572. doi:10.2305/IUCN.UK.2016-3.RLTS.T19646A22275131.en.
  16. ^ a b c Nowak, p. 1442
  17. ^ a b Aplin, K.; Molur, S. (2017) [errata version of 2016 assessment]. "Rhizomys pruinosus". IUCN Red List of Threatened Species. 2016 e.T19645A115152385. doi:10.2305/IUCN.UK.2016-3.RLTS.T19645A22274964.en.
  18. ^ a b Aplin, K.; Lunde, D. (2017) [errata version of 2016 assessment]. "Rhizomys sumatrensis". IUCN Red List of Threatened Species. 2016 e.T19647A115152803. doi:10.2305/IUCN.UK.2016-3.RLTS.T19647A22275342.en.
  19. ^ a b Lavrenchenko, L.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Tachyoryctes macrocephalus". IUCN Red List of Threatened Species. 2016 e.T21293A115161321. doi:10.2305/IUCN.UK.2016-3.RLTS.T21293A22276163.en.
  20. ^ a b Nowak, p. 1445
  21. ^ a b Cassola, F. (2017). "Tachyoryctes splendens". IUCN Red List of Threatened Species. 2017 e.T21299A22275532. doi:10.2305/IUCN.UK.2017-2.RLTS.T21299A22275532.en.
  22. ^ a b Arslan, A.; Gazzard, A.; Matur, F.; Sozen, M. (2023). "Nannospalax xanthodon". IUCN Red List of Threatened Species. 2023 e.T14327A22276510. doi:10.2305/IUCN.UK.2023-1.RLTS.T14327A22276510.en.
  23. ^ a b c d e f g h i j k Nowak, p. 1427
  24. ^ a b Rusin, M. (2024). "Nannospalax leucodon". IUCN Red List of Threatened Species. 2024 e.T14328A221788646. doi:10.2305/IUCN.UK.2024-2.RLTS.T14328A221788646.en.
  25. ^ a b Lövy, M.; Gazzard, A. (2023). "Nannospalax ehrenbergi". IUCN Red List of Threatened Species. 2023 e.T14326A22276839. doi:10.2305/IUCN.UK.2023-1.RLTS.T14326A22276839.en.
  26. ^ a b Rusin, M. (2024). "Spalax graecus". IUCN Red List of Threatened Species. 2024 e.T97249856A217345371. doi:10.2305/IUCN.UK.2024-2.RLTS.T97249856A217345371.en.
  27. ^ a b Kennerley, R.; Formozov, N.; Sheftel, B. (2016). "Spalax giganteus". IUCN Red List of Threatened Species. 2016 e.T20429A2772339. doi:10.2305/IUCN.UK.2016-2.RLTS.T20429A2772339.en.
  28. ^ a b Rusin, M. (2024). "Spalax microphthalmus". IUCN Red List of Threatened Species. 2024 e.T20430A221789991. doi:10.2305/IUCN.UK.2024-2.RLTS.T20430A221789991.en.
  29. ^ a b Rusin, M.; Gazzard, A. (2025). "Spalax uralensis". IUCN Red List of Threatened Species. 2025 e.T136581A281170219.
  30. ^ a b Németh, A.; Csorba, G.; Hegyeli, Z. (2024). "Spalax antiquus". IUCN Red List of Threatened Species. 2024 e.T97250195A221786525. doi:10.2305/IUCN.UK.2024-2.RLTS.T97250195A221786525.en.
  31. ^ a b Hegyeli, Z.; Csorba, G.; Németh, A. (2024). "Spalax istricus". IUCN Red List of Threatened Species. 2024 e.T97250154A217345946. doi:10.2305/IUCN.UK.2024-2.RLTS.T97250154A217345946.en.
  32. ^ a b Rusin, M. (2024). "Spalax zemni". IUCN Red List of Threatened Species. 2024 e.T42655A91863646. doi:10.2305/IUCN.UK.2024-2.RLTS.T42655A91863646.en.
  33. ^ a b Rusin, M. (2025) [errata version of 2024 assessment]. "Spalax arenarius". IUCN Red List of Threatened Species. 2024 e.T20428A270136276. doi:10.2305/IUCN.UK.2024-2.RLTS.T20428A270136276.en.

Sources

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